After a 1000-year war between the Autobots and the Decepticons, Optimus and Co. are finally able to rest. They build the Dome of Formation, and let Humanity rebuild itself. 800 years later, however, just as things started getting easier, a new problem arose: Callbots. These kleptomaniacs and murderers have one thing in mind: clear out all humans and drain Earth of all its natural wealth. They can share thoughts through the Mother System, which Optimus and his new friend Cal have to destroy to save the planet.