On the eve of a major anniversary, the Fantastic Four invite their friends and allies to the Baxter Building for a huge celebration. Old photographs, newspaper clippings, and souvenirs bring back memories of their earliest adventures. However, the celebration takes an unexpected turn when the family discovers that their arch-nemesis Dr. Doom has been secretly manipulating events throughout their history. To stop Doom, the Four must revisit the past and uncover the truth behind one of their forgotten adventures.