Iggy and Spike realize they forgot Daisy's rock, a meteorite fragment that Koopa wants to obtain to merge his world with the human world. Daisy turns out to be the long-lost princess of the other dimension. When Koopa overthrew Daisy's father as king and devolved him into fungus, her mother the queen took her to Brooklyn. The portal was then sealed, but Scapelli's men inadvertently reopened the portal when they blasted the cave. Koopa sends his cousins to find Daisy and the rock to merge the dimensions and make him dictator of both worlds. However, after Koopa subjects them to an experiment to increase their intelligence, they realize Koopa's evil intentions and side with the Mario Bros. Daisy is taken to Koopa Tower, where she meets Yoshi. Koopa informs Daisy that she descended from the dinosaurs, believing only she can merge the worlds because of her royal heritage. The Mario Bros. rescue Daisy, aided by Toad, a good-natured guitarist who was devolved into a Goomba, a semi-humanoid dinosaur, as punishment for a protest.
