Colonel J. Wesley McCullough was a United States Army Special Forces officer and the Commanding Officer of Alpha-Omega, his unit hunting down the apes. Because of his ruthless nature and the drastic actions he took to preserve human purity and supremacy, he was feared by the remaining army leadership. During the course of War for the Planet of the Apes, it is apparent he's not only at war with the Ape Colony led by Caesar, but also against the remaining U.S. Army.