Darwinopterus (meaning "Darwin's wing") is a genus of pterosaur, discovered in China and named after biologist Charles Darwin.[1] Between 30 and 40 fossil specimens have been identified,[2] all collected from the Tiaojishan Formation, which dates to the middle Jurassic period, 160.89–160.25 Ma ago.[3][4] The type species, D. modularis, was described in February 2010.[1] D. modularis was the first known pterosaur to display features of both long-tailed (rhamphorhynchoid) and short-tailed (pterodactyloid) pterosaurs, and was described as a transitional fossil between the two groups.[5] Three additional species, D. camposi, D. linglongtaensis, and D. robustodens, were described from the same fossil beds in February 2025,[6] December 2010,[7] and June 2011, respectively.[8]