Archelon is an extinct marine turtle from the Late Cretaceous, and is the largest turtle ever to have been documented, with the biggest specimen measuring 460 cm (15 ft) from head to tail, 400 cm (13 ft) from flipper to flipper, and 2,200 kg (4,900 lb) in weight. It is known only from the Dakota Pierre Shale and has one species, A. ischyros. In the past, the genus also contained A. marshii and A. copei, though these have been reassigned to Protostega and Microstega, respectively. The genus was named in 1895 by American paleontologist George Reber Wieland based on a skeleton from South Dakota, who placed it into the extinct family Protostegidae. The leatherback sea turtle (Dermochelys coriacea) was once thought to be its closest living relative, but now, Protostegidae is thought to be a completely separate lineage from any living sea turtle. Archelon had a leathery carapace instead of the hard shell seen in sea turtles. The carapace may have featured a row of small ridges, each peaking at 2.5 or 5 cm (1 or 2 in) in height. It had an especially hooked beak and its jaws were adept at crushing, so it probably ate hard-shelled crustaceans and mollusks while slowly moving over the seafloor. However, its beak may have been adapted for shearing flesh, and Archelon was likely able to produce powerful strokes necessary for open-ocean travel. It inhabited the northern Western Interior Seaway, a mild to cool area dominated by plesiosaurs, hesperornithiform seabirds, and mosasaurs. It may have gone extinct due to the shrinking of the seaway, increased egg and hatchling predation, and cooling climate.