Following the Dimension Tide assault, Godzilla appears in an alternate Earth conquered by the Millennians. This dying planet, once a parallel Earth, was consumed by their greed for plasma energy. The world's guardian monsters are controlled by the Millennian empire, but when Godzilla arrives, they send in the once guardian of the universe, Gamera to attack, along with Mothra, Varan, Anguirus, and Baragon. A small group of rebels infiltrate the kingdom and try to set the guardians free. This prompts the mysterious Millennian leader to unleash their ultimate weapon, King Ghidorah.
