Last time, the monsters were defeated. Rodan lay fallen, and Godzilla claimed his place as King of the Monsters, seated upon a throne of scorched rubble and ruin. But his reign is finally challenged by the one true guardian destined to face him: Gamera. Meanwhile, humanity stands on the brink of extinction. Deep within an underground shelter, scientists propose recreating the Oxygen Destroyer using modern technology. Ogata is reluctant at first, unwilling to betray the memory of his old friend, Dr. Serizawa, and the sacrifice he made so long ago. But with no other hope left, he agrees. If humanity as well as other kaiju life are to survive, Godzilla must be destroyed once and for all. As preparations for the weapon begin, Gamera, with Rodan fighting at his side, confronts his long-awaited enemy. In the ruins of a dying world, Gamera and Godzilla clash in a final battle to the death.