Zoé Lee is an American teenage girl from New York City and a student in Ms. Mendeleiev's class at Collège Françoise Dupont. She is also Chloé Bourgeois's sister and André Bourgeois's stepdaughter. She attended a boarding school when she was in New York, where she once lived with her mother. Her father is American and her mother is French-American. In "Sole Crusher", after being tired of hiding her true self in order to please her sister, Zoé is akumatized by Shadow Moth into Sole Crusher, a supervillain who can absorb people and becomes larger by stomping on or kicking them. She also receives a Magical Charm from Ladybug that prevents her from being akumatized again. In "Queen Banana", after Chloé is akumatized into the titular villain, Zoé receives the Bee Miraculous from Ladybug. When the Miraculous is inhabited by Pollen, Zoé becomes Vesperia, a bee/wasp-themed superhero who helps Ladybug and Cat Noir during particularly difficult missions in the fourth season.