Desmond, William, Rebecca and Shaun find the Grand Temple of the First Civilization in a cave in New York and access it using the Apple of Eden. After partially activating the temple, Juno begins to communicate with Desmond. Desmond is placed in the Animus, where Juno's influence causes him to experience the life of his ancestor, Haytham Kenway.The perspective then shifts to the childhood years of Ratonhnhaké:ton, Haytham and Ziio's son. In 1760, his mother dies during an attack on his village, which Ratonhnhaké:ton assumes was led by Lee. Years later, the village elder informs Ratonhnhaké:ton that it is their tribe's duty to prevent the Temple's discovery, and gives him a Sphere which allows Juno to communicate with him. She leads Ratonhnhaké:ton to retired Assassin Achilles Davenport, whose brotherhood collapsed years ago, and he reluctantly agrees to train him. After completing his training, Achilles suggests Ratonhnhaké:ton rename himself "Connor" to move more freely throughout the Colonies.
